Mobile element type:  Transposon
Name:  Tn1696
Synonyms: 
Accession:  Tn1696-U12338.3
Family:  Tn3
Group:  Tn21
First isolate:  Yes
Partial:  ND
Evidence of transposition:  ND

Host Organism:  Pseudomonas aeuroginosa
Date of Isolation: 
Country:  Spain
Molecular Source:  plasmid R1033

Name Coordinates Direction Length Sequence
IRL 1-38 + 38 GGGGTCGTCT CAGAATTCGG AAAATAAAGC ACGCTAAG
IRR 16281-16318 - 38 GGGGTCGTCT CAGAAAACGG AAAATAAAGC ACGCTAAG

Name Associated TE Coordinates Orientation Length Sequence
res_site_I Tn1696 4278-4308 + 31 CGTCAGATTG AGGCATACCC TAACTTGATG T
attC-qacE2 3'-end In4 6118-6123 + 6 TTAGAT
attC qacH_sul1 In4 6718-6751 , 8029-8035 + 41 CCGCTAGCGG GCGGCCGGAA GGTGAATGCT AGGCATCTAA C
attC qacH_sul1 core In4 6718-6751 + 34 CCGCTAGCGG GCGGCCGGAA GGTGAATGCT AGGC
attC-qacE2 3'-end In4 6755-6760 + 6 ATCTAA
attC-cmlA6 In4 8035-8098 , 9578-9583 + 70 CGCCTGAGCT CAGCCGACCG AAACCGCGTA GCGGTTTTGG GTCGGCTGCA GCGATTTGTT GGGCGCCCAA
attC cmlA6 core In4 8035-8098 + 64 CGCCTGAGCT CAGCCGACCG AAACCGCGTA GCGGTTTTGG GTCGGCTGCA GCGATTTGTT GGGC
attC-cmlA6 3'-end In4 9578-9583 + 6 GCCCAA
attC-aadA3 In4 9584-9637 , 10434-10439 + 60 CGCCGGAGTT AAGCCGCCGC GCGTAGCGCG GTCGGCTTGA ACGAATTGTT AGACGTCTAA
attC aadA3 core In4 9584-9637 + 54 CGCCGGAGTT AAGCCGCCGC GCGTAGCGCG GTCGGCTTGA ACGAATTGTT AGAC
attC-DUF1010 In4 10440-10493 , 10696-10701 + 60 CGGCCAAGGT AAGCGGCCCG CAGAATGCGG GTCCGCTTGA CCGCAGAGTT AGACGACTAA
attC DUF1010 core In4 10440-10493 + 54 CGGCCAAGGT AAGCGGCCCG CAGAATGCGG GTCCGCTTGA CCGCAGAGTT AGAC
attC-DUF1010 3'-end In4 10696-10701 + 6 GACTAA
attC-AAC(3)-Ia In4 10702-10804 , 11273-11278 + 109 CGCCTGAGTT AAGCCGGAGC GCTTTGCGGC CGCGGCGTTG TGACAATTTA CCGAACAACT CCGCGGCCGG GAAGCCGATC TCGGCTTGAA CGAATTGTTA GGTACCTAA
attC AAC(3)-Ia core In4 10702-10804 + 103 CGCCTGAGTT AAGCCGGAGC GCTTTGCGGC CGCGGCGTTG TGACAATTTA CCGAACAACT CCGCGGCCGG GAAGCCGATC TCGGCTTGAA CGAATTGTTA GGT
attC-AAC(3)-Ia 3'-end In4 11273-11278 + 6 ACCTAA
res_site_II Tn1696 12664-12698 + 35 TGTCAGAATA GAGTTAAATT TCCTATTGAT TGACA
res_site_III Tn1696 12701-12732 + 32 TTCCGTCAAA GGTAATAGAT TTCATCCTGA CA

ORF Summary
Name Associated TE Coordinates Orientation Class Subclass
merR Tn1696 34-489 - Passenger Gene Heavy Metal Resistance
merT Tn1696 561-926 + Passenger Gene Heavy Metal Resistance
merP Tn1696 942-1217 + Passenger Gene Heavy Metal Resistance
merC Tn1696 1245-1670 + Passenger Gene Heavy Metal Resistance
merA Tn1696 1709-3394 + Passenger Gene Heavy Metal Resistance
merD Tn1696 3412-3777 + Passenger Gene Heavy Metal Resistance
merE Tn1696 3774-4010 + Passenger Gene Heavy Metal Resistance
tnp IS6100 4847-5611 - Transposase
WP_001424634.1 In4 5807-6094 - Passenger Gene Hypothetical
GNAT_fam In4 6118-6618 - Passenger Gene Antibiotic Resistance
sul1 (ARO:3000410) In4 6746-7585 - Passenger Gene Antibiotic Resistance
qacEdelta1 (ARO:3005010) In4 7579-7926 - Passenger Gene Antibiotic Resistance
cmlA6 (ARO:3002696) In4 8118-9377 - Passenger Gene Antibiotic Resistance
aadA3 (ARO:3002603) In4 9639-10430 - Passenger Gene Antibiotic Resistance
DUF1010 family protein In4 10436-10681 - Passenger Gene Other
AAC(3)-Ia (ARO:3002528) In4 10799-11263 - Passenger Gene Antibiotic Resistance
intI1 In4 11434-12447 + Integron Integrase Class 1
tnpR Tn1696 12753-13310 + Accessory Gene Resolvase
tnpA Tn1696 13313-16285 + Transposase
ORF Details
Gene Name Protein Name Associated TE Gene Length Coordinates Strand
merR MerR Tn1696 151 34-489 -
Class: Passenger Gene
Subclass: Heavy Metal Resistance
Target: Mercury
Protein Sequence: MQINFENLTI GVFAKAAGVN VETIRFYQRK GLLPEPDKPY GSIRRYGEAD VTRVRFVKSA QRLGFSLDEI AELLRLEDGT HCEEASGLAE HKLKDVREKM ADLARMEAVL SELVCACHAR KGNVSCPLIA SLQDGTKLAA SARGSHGVTT P

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
merT MerT Tn1696 121 561-926 +
Class: Passenger Gene
Subclass: Heavy Metal Resistance
Target: Mercury
Protein Sequence: MSEPQKSEPQ NGRGALFAGG LAAILASACC LGPLVLIALG FSGAWIGNLT VLEPYRPIFI GAALVALFFA WRRIYRPAQA CKPGEVCAIP QVRATYKLIF WIVAALVLVS LGFPYVMPFF Y

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
merP MerP Tn1696 91 942-1217 +
Class: Passenger Gene
Subclass: Heavy Metal Resistance
Target: Mercury
Protein Sequence: MKKLFAALAL AAVVAPVWAA TQTVTLSVPG MTCASCPITV KHALSKVEGV SKTDVSFDKR QAVVTFDDAK TNVQKLTKAT EDAGYPSSLK R

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
merC MerC Tn1696 141 1245-1670 +
Class: Passenger Gene
Subclass: Heavy Metal Resistance
Target: Mercury
Protein Sequence: MGLITRIAGK TGALGSVVSA MGCAACFPAI ASFGAAIGLG FLSQYEGLFI GILLPMFAGI ALLANAIAWL NHRQWRRTAL GTIGPILVLA AVFLMRAYGW QSGGLLYVGL ALMVGVSVWD FISPAHRRCG PDSCELPEQR G

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
merA MerA Tn1696 561 1709-3394 +
Class: Passenger Gene
Subclass: Heavy Metal Resistance
Target: Mercury
Protein Sequence: MTTLKITGMT CDSCAAHVKE ALEKVPGVQS ALVSYPKGTA QLAIEAGTSS DALTTAVAGL GYEATLADAP PTDNRAGLLD KMRGWIGAAD KPSGNERPLQ VVVIGSGGAA MAAALKAVEQ GAQVTLIERG TIGGTCVNVG CVPSKIMIRA AHIAHLRRES PFDGGMPPTP PTILRERLLA QQQARVEELR HAKYEGILDG NSAITVLHGE ARFKDDQSLI VSLNEGGERV VMFDRCLVAT GASPAVPPIP GLKESPYWTS TEALASDTIP ERLAVIGSSV VALELAQAFA RLGSKVTALA RNTLFFREDP AIGEAVTAAF RAEGIEVLEH TQASQVAHMD GEFVLTTTHG ELRADKLLVA TGRTPNTRSL ALEAAGVAVN AQGAIVIDKG MRTSSPNIYA AGDCTDQPQF VYVAAAAGTR AAINMTGGDA ALDLTAMPAV VFTDPQVATV GYSEAEAHHD GIETDSRLLT LDNVPRALAN FDTRGFIKLV IEEGSGRLIG VQAVAPEAGE LIQTAVLAIR NRMTVQELAD QLFPYLTMVE GLKLAAQTFS KDVKQLSCCA G

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
merD MerD Tn1696 121 3412-3777 +
Class: Passenger Gene
Subclass: Heavy Metal Resistance
Target: Mercury
Protein Sequence: MSAYTVSRLA LDAGVSVHIV RDYLLRGLLR PVAYTTGGYG LFDDTALQRL RFVRAAFEAG IGLDALARLC RALDAADGDG ASAQLAVLRQ LVERRREALA SLEMQLAAMP TEPAQHAESL P

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
merE MerE Tn1696 78 3774-4010 +
Class: Passenger Gene
Subclass: Heavy Metal Resistance
Target: Mercury
Protein Sequence: MNSPEHLPSE THKPITGYLW GALAVLTCPC HLPILAIVLA GTTAGAFIGE HWGIAALTLT GLFVLSVTRL LRAFKGRS

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
tnp Tnp IS6100 254 4847-5611 -
Class: Transposase
Transposase Chemistry: DDE
Protein Sequence: MTDFKWRHFQ GDVILWAVRW YCRYPISYRD LEEMLAERGI SVDHTTIYRW VQCYAPEMEK RLRWFWRRGF DPSWRLDETY VKVRGKWTYL YRAVDKRGDT IDFYLSPTRS AKAAKRFLGK ALRGLKHWEK PATLNTDKAP SYGAAITELK REGKLDRETA HRQVKYLNNV IEADHGKLKI LIKPVRGFKS IPTAYATIKG FEVMRALRKG QARPWCLQPG IRGEVRLVER AFGIGPSALT EAMGMLNHHF AAAA

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
WP_001424634.1 WP_001424634.1 In4 95 5807-6094 -
Class: Passenger Gene
Subclass: Hypothetical
Protein Sequence: MSGWDGPVAS RSLAVRPRCL GCARGCRYPP LPSSTLDNAA AVIAADARAG LFGPVRLARP SACLGEISGG RINGGASPPF VRVKAEDSSL KNIRL

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
GNAT_fam GNAT_fam In4 166 6118-6618 -
Class: Passenger Gene
Subclass: Antibiotic Resistance
Sequence Family: GNAT
Target: aminoglycoside antibiotic (ARO:0000016)
Protein Sequence: MDSEEPPNVR VACSGDIDEV VRLMHDAAAW MSAKGTPAWD VARIDRTFAE TFVLRSELLV ASCSDGIVGC CTLSAEDPEF WPDALKGEAA YLHKLAVRRT HAGRGVSSAL IEACRHAART QGCAKLRLDC HPNLRGLYER LGFTHVDTFN PGWDPTFIAE RLELEI

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
sul1 (ARO:3000410) Sul1 In4 279 6746-7585 -
Class: Passenger Gene
Subclass: Antibiotic Resistance
Function: antibiotic target replacement (ARO:0001002)
Sequence Family: sulfonamide resistant sul (ARO:3004238)
Transposase Chemistry: dihydropteroate synthase
Target: sulfonamide antibiotic (ARO:3000282)||sulfone antibiotic (ARO:3003401)
Comment: perfect match to reference sequence for ARO:3000410
Protein Sequence: MVTVFGILNL TEDSFFDESR RLDPAGAVTA AIEMLRVGSD VVDVGPAASH PDARPVSPAD EIRRIAPLLD ALSDQMHRVS IDSFQPETQR YALKRGVGYL NDIQGFPDPA LYPDIAEADC RLVVMHSAQR DGIATRTGHL RPEDALDEIV RFFEARVSAL RRSGVAADRL ILDPGMGFFL SPAPETSLHV LSNLQKLKSA LGLPLLVSVS RKSFLGATVG LPVKDLGPAS LAAELHAIGN GADYVRTHAP GDLRSAITFS ETLAKFRSRD ARDRGLDHA

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
qacEdelta1 (ARO:3005010) QacEdelta1 In4 115 7579-7926 -
Class: Passenger Gene
Subclass: Antibiotic Resistance
Function: antibiotic efflux (ARO:0010000)
Sequence Family: major facilitator superfamily (MFS) antibiotic efflux pump (ARO:0010002)
Target: disinfecting agents and antiseptics(ARO:3005386)
Comment: subunit of the qac multidrug efflux pump||perfect match to reference sequence for ARO:3005010 (bitscore:219)
Protein Sequence: MKGWLFLVIA IVGEVIATSA LKSSEGFTKL APSAVVIIGY GIAFYFLSLV LKSIPVGVAY AVWSGLGVVI ITAIAWLLHG QKLDAWGFVG MGLIIAAFLL ARSPSWKSLR RPTPW

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
cmlA6 (ARO:3002696) CmlA6 In4 419 8118-9377 -
Class: Passenger Gene
Subclass: Antibiotic Resistance
Function: antibiotic efflux (ARO:0010000)
Sequence Family: major facilitator superfamily (MFS) antibiotic efflux pump (ARO:0010002)
Target: phenicol antibiotic (ARO:3000387)
Comment: strict match to reference sequence for ARO:3002696 (bitscore: 819)
Protein Sequence: MSSKNFSWRY SLAATVLLLS PFDLLASLGM DMYLPAVPFM PNALGTTAST IQLTLTTYLV MIGAGQLLFG PLSDRLGRRP VLLGGGLAYV VASMGLALTS SAEVFLGLRI LQACGASACL VSTFATVRDI YAGREESNVI YGILGSMLAM VPAVGPLLGA LVDMWLGWRA IFAFLGLGMI AASAAAWRFW PETRVQRVAG LQWSQLLLPV KCLNFWLYTL CYAAGMGSFF VFFSIAPGLM MGRQGVSQLG FSLLFATVAI AMVFTARFMG RVIPKWGSPS VLRMGMGCLI AGAVLLAITE IWALQSVLGF IAPMWLVGIG VATAVSVAPN GALRGFDHVA GTVTAVYFCL GGVLLGSIGT LIISLLPRNT AWPVVVYCLT LATVVLGLSC VSRVKGSRGQ GEHDVVALQS AESTSNPNR

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
aadA3 (ARO:3002603) AadA3 In4 263 9639-10430 -
Class: Passenger Gene
Subclass: Antibiotic Resistance
Function: antibiotic inactivation (ARO:0001004)
Sequence Family: ANT(3'')(ARO:3004275)
Target: aminoglycoside antibiotic (ARO:0000016)
Comment: strict match to reference sequence for ARO:3002603 (bitscore: 522)
Protein Sequence: MRVAVTIEIS NQLSEVLSVI ERHLESTLLA VHLYGSAVDG GLKPYSDIDL LVTVAVKLDE TTRRALLNDL MEASAFPGES ETLRAIEVTL VVHDDIIPWR YPAKRELQFG EWQRNDILAG IFEPAMIDID LAILLTKARE HSVALVGPAA EEFFDPVPEQ DLFEALRETL KLWNSQPDWA GDERNVVLTL SRIWYSAITG KIAPKDVAAD WAIKRLPAQY QPVLLEAKQA YLGQKEDHLA SRADHLEEFI RFVKGEIIKS VGK

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
DUF1010 family protein DUF1010 family protein In4 81 10436-10681 -
Class: Passenger Gene
Subclass: Other
Sequence Family: DUF1010 (Pfam:PF06231)
Comment: BLAST: WP_032492246.1
Protein Sequence: MRAATYFSAP PCSKGFLAFW ASSACAASAT SYPSCSAAPL SWSSGFSWAA PFFKAGRSLL AFGSNSAVKR TRILRAAYLG R

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
AAC(3)-Ia (ARO:3002528) AAC(3)-Ia In4 154 10799-11263 -
Class: Passenger Gene
Subclass: Antibiotic Resistance
Function: antibiotic inactivation (ARO:0001004)
Sequence Family: AAC(3) (ARO:3000322)
Target: aminoglycoside antibiotic (ARO:0000016)
Comment: perfect match to reference sequence for ARO:3002528||Synonyms: AAC(3)-Ia C, aacC1, aacC-A1
Protein Sequence: MGIIRTCRLG PDQVKSMRAA LDLFGREFGD VATYSQHQPD SDYLGNLLRS KTFIALAAFD QEAVVGALAA YVLPRFEQPR SEIYIYDLAV SGEHRRQGIA TALINLLKHE ANALGAYVIY VQADYGDDPA VALYTKLGIR EEVMHFDIDP STAT

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
intI1 IntI1 In4 337 11434-12447 +
Class: Integron Integrase
Subclass: Class 1
Sequence Family: Class 1 Integron Tyrosine Integrase
Transposase Chemistry: Tyrosine
Protein Sequence: MKTATAPLPP LRSVKVLDQL RERIRYLHYS LRTEQAYVNW VRAFIRFHGV RHPATLGSSE VEAFLSWLAN ERKVSVSTHR QALAALLFFY GKVLCTDLPW LQEIGRPRPS RRLPVVLTPD EVVRILGFLE GEHRLFAQLL YGTGMRISEG LQLRVKDLDF DHGTIIVREG KGSKDRALML PESLAPTLRE QLSRARAWWL KDQAEGRSGV ALPDALERKY PRAGHSWPWF WVFAQHTHST DPRSGVVRRH HMYDQTFQRA FKRAVEQAGI TKPATPHTLR HSFATALLRS GYDIRTVQDL LGHSDVSTTM IYTHVLKVGG AGVRSPLDAL PPLTSER

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
tnpR TnpR Tn1696 185 12753-13310 +
Class: Accessory Gene
Subclass: Resolvase
Sequence Family: Serine Site-SpecificRecombinase
Transposase Chemistry: Serine
Protein Sequence: MQGQRIGYVR VSSFDQNPER QLEGVQVARV FTDKASGKDT QRPELERLLA FVREGDTVVV HSMDRLARNL DDLRRIVQGL TQRGVRMEFV KEGLKFTGED SPMANLMLSV MGAFAEFERA LIRERQREGI VLAKQRGAYR GRKKSLNSEQ IAELKRRVAA GDQKTLVARD FGISRETLYQ YLRED

Gene Name Protein Name Associated TE Gene Length Coordinates Strand
tnpA TnpA Tn1696 990 13313-16285 +
Class: Transposase
Transposase Chemistry: DDE
Protein Sequence: MPRRSILSAT ERESLLALPD AKDELIRHYT FNETDLSVIR QRRGAANRLG FAVQLCYLRF PGTFLGVDEP PFPPLLRMVA AQLKMPVESW SEYGQREQTR REHLVELQTV FGFKPFTMSH YRQAVHTLTE LALQTDKGIV LASALVENLR RQSIILPAMN AIERASAEAI TRANRRIYAA LTDSLLSPHR QRLDELLKRK DGSKVTWLAW LRQSPAKPNS RHMLEHIERL KSWQALDLPA GIERQVHQNR LLKIAREGGQ MTPADLAKFE VQRRYATLVA LAIEGMATVT DEIIDLHDRI IGKLFNAAKN KHQQQFQASG KAINDKVRMY GRIGQALIEA KQSGSDPFAA IEAVMPWDTF AASVTEAQTL ARPADFDFLH HIGESYATLR RYAPQFLGVL KLRAAPAAKG VLDAIDMLRG MNSDSARKVP ADAPTAFIKP RWAKLVLTDD GIDRRYYELC ALSELKNALR SGDVWVQGSR QFKDFDEYLV PVEKFATLKL ASELPLAVAT DCDQYLHDRL ELLEAQLATV NRMAAANDLP DAIITTASGL KITPLDAAVP DAAQAMIDQT AMLLPHLKIT ELLMEVDEWT GFTRHFTHLK TSDTAKDKTL LLTTILADAI NLGLTKMAES CPGTTYAKLS WLQAWHIRDE TYSTALAELV NAQFRQPFAG NWGDGTTSSS DGQNFRTGSK AESTGHINPK YGSSPGRTFY THISDQYAPF SAKVVNVGIR DSTYVLDGLL YHESDLRIEE HYTDTAGFTD HVFGLMHLLG FRFAPRIRDL GETKLFIPKG DAAYDALKPM ISSDRLNIKQ IRAHWDEILR LATSIKQGTV TASLMLRKLG SYPRQNGLAV ALRELGRIER TLFILDWLQS VELRRRVHAG LNKGEARNAL ARAVFFYRLG EIRDRSFEQQ RYRASGLNLV TAAIVLWNTV YLERATSALR GNGTALDDTL LQYLSPLGWE HINLTGDYLW RSSAKVGAGK FRPLRPLPPA

TnCentral Accession TE Name Type Coordinates Strand Length
In4-U12338.3 In4 integron 4320-12649 + 8330
IS6100-U12338.3 IS6100 insertion sequence 4793-5672 + 880

Name Associated TE Coordinates Direction Length Sequence
IRt In4 4320-4352 + 33 TGTCATTTTC AGAAGACGAC TGCACCAGTT GAT
repeat t1 In4 4328-4346 + 19 TCAGAAGACG ACTGCACCA
repeat t2 In4 4368-4386 - 19 TCAGGAGCTG GCTGCACAA
repeat t3 In4 4397-4416 + 20 TCAGAAGTGA TCTGCACCAA
repeat t4 In4 4429-4447 + 19 TCAATACTCG TGTGCACCA
IRR IS6100 4793-4806 + 14 GGCTCTGTTG CAAA
IRL IS6100 5659-5672 - 14 GGCTCTGTTG CAAA
repeat t3 In4 5699-5718 - 20 TCAGAAGTGA TCTGCACCAA
repeat t2 In4 5729-5747 + 19 TCAGGAGCTG GCTGCACAA
IRt In4 5763-5795 - 33 TGTCATTTTC AGAAGACGAC TGCACCAGTT GAT
repeat t1 In4 5769-5787 - 19 TCAGAAGACG ACTGCACCA
repeat i4 In4 12530-12548 - 19 TCAGCGGACG CAGGGAGGA
repeat i3 In4 12558-12576 - 19 TCAGGCAACG ACGGGCTGC
repeat i2 In4 12600-12618 - 19 TCAGAAGCCG ACTGCACTA
IRi In4 12617-12649 - 33 TGTCGTTTTC AGAAGACGGC TGCACTGAAC GTC
repeat i1 In4 12623-12641 - 19 TCAGAAGACG GCTGCACTG

1.  Wohlleben W, Arnold W, Bissonnette L, Pelletier A, Tanguay A, Roy PH, Gamboa GC, Barry GF, Aubert E, Davies J, et al..  On the evolution of Tn21-like multiresistance transposons: sequence analysis of the gene (aacC1) for gentamicin acetyltransferase-3-I(AAC(3)-I), another member of the Tn21-based expression cassette..  Mol Gen Genet. 1989 Jun;217(2-3):202-8. doi: 10.1007/BF02464882.  PubMed ID: 2549372 .
2.  Bissonnette L, Champetier S, Buisson JP, Roy PH.  Characterization of the nonenzymatic chloramphenicol resistance (cmlA) gene of the In4 integron of Tn1696: similarity of the product to transmembrane transport proteins..  J Bacteriol. 1991 Jul;173(14):4493-502. doi: 10.1128/jb.173.14.4493-4502.1991.  PubMed ID: 1648560 .
3.  Stokes HW, Hall RM.  Sequence analysis of the inducible chloramphenicol resistance determinant in the Tn1696 integron suggests regulation by translational attenuation..  Plasmid. 1991 Jul;26(1):10-9. doi: 10.1016/0147-619x(91)90032-r.  PubMed ID: 1658833 .
4.  Collis CM, Hall RM.  Site-specific deletion and rearrangement of integron insert genes catalyzed by the integron DNA integrase..  J Bacteriol. 1992 Mar;174(5):1574-85. doi: 10.1128/jb.174.5.1574-1585.1992.  PubMed ID: 1311297 .
5.  Hall RM, Brown HJ, Brookes DE, Stokes HW.  Integrons found in different locations have identical 5' ends but variable 3' ends..  J Bacteriol. 1994 Oct;176(20):6286-94. doi: 10.1128/jb.176.20.6286-6294.1994.  PubMed ID: 7929000 .
6.  Partridge SR, Brown HJ, Stokes HW, Hall RM.  Transposons Tn1696 and Tn21 and their integrons In4 and In2 have independent origins..  Antimicrob Agents Chemother. 2001 Apr;45(4):1263-70. doi: 10.1128/AAC.45.4.1263-1270.2001.  PubMed ID: 11257044 .
7.  Smith DI, Lus RG, Rubio Calvo MC, Datta N, Jacob AE, Hedges RW.  Third type of plasmid conferring gentamicin resistance in Pseudomonas aeruginosa..  Antimicrob Agents Chemother. 1975 Sep;8(3):227-30. doi: 10.1128/AAC.8.3.227.  PubMed ID: 810080 .